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Phosphorylated TrkB Tyr 516 through the interaction with scaffold protein Shc (Src homologous and collagen-like) mediates the activation of PI3K (Phosphatidylinositol 3-kinase) via Grb2 (Growth factor receptor-bound protein 2) and Gab1 (Grb2-associated binder-1) proteins, which in turn leads to Akt (Rac-alpha serine/threonine-protein kinase) activation, promotion of Akt-dependent neurotrophin survival effects and increased protein translation (regulated by mTOR-p70S6 kinase)

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